The Omnivore's Dilemma: A Natural History of Four Meals

by Michael Pollan

The C-4 trick represents an important economy for a plant, giving it an advantage, especially in areas where water is scarce and temperatures high. In order to gather carbon atoms from the air, a plant has to open its stomata, the microscopic orifices in the leaves through which plants both take in and exhaust gases. Every time a stoma opens to admit carbon dioxide precious molecules of water escape. It’s as though every time you opened your mouth to eat you lost a quantity of blood. Ideally, you would open your mouth as seldom as possible, ingesting as much food as you could with every bite. This is essentially what a C-4 plant does. By recruiting extra atoms of carbon during each instance of photosynthesis, the corn plant is able to limit its loss of water and “fix”—that is, take from the atmosphere and link in a useful molecule—significantly more carbon than other plants.