O’Neill’s A Hierarchical Concept of Ecosystems. O’Neill and his co-authors noted that ecosystems could be better understood by observing the rates of change of different components. Hummingbirds and flowers are quick, redwood trees slow, and whole redwood forests even slower. Most interaction is within the same pace level—hummingbirds and flowers pay attention to each other, oblivious to redwoods, who are oblivious to them. Meanwhile the forest is attentive to climate change but not to the hasty fate of individual trees. The insight is this: “The dynamics of the system will be dominated by the
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