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Different mutations affecting male behaviors or traits could appear in different places—say, longer tail feathers in one population and orange color in another—and sexual selection might then drive the populations in different directions. Eventually, females in one population would prefer long-tailed males, and females in the other, orange males. If the two populations later encountered each other, their mating preferences would prevent them from mixing genes, and they would be considered different species.
Why Evolution Is True
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