When Charles Darwin described flowers whose nectar reservoirs could be reached only through a narrow two-inch flower tube, he famously predicted that there must be some undiscovered insect with an equally long tongue. It was many years after Darwin’s death before such insects were discovered. The meganosed fly (Moegistorhynchus longirostris) of southern Africa, outdoing its literary counterpart Pinocchio, has a noselike proboscis that is actually the longest mouthpart—a tonguelike drinking tube—of any known fly

