The anatomical alterations in the slug’s relatively simple memory circuits were extensive. In one case, the researchers found that, before a long-term memory was consolidated, a particular sensory neuron had some thirteen hundred synaptic connections to about twenty-five other neurons. Only about forty percent of those connections were active—in other words, sending signals through the production of neurotransmitters. After the long-term memory had been formed, the number of synaptic connections had more than doubled, to about twenty-seven hundred, and the proportion that were active had
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