‘And it must be hardest on Rani, knowing that the young ones depend on her. There she is, perfectly adapted to her environment, perfectly at home in it – and then things begin to change, so that all those years of learning become useless, the places you know best can’t sustain you any more and you’ve got to find new hunting grounds. Rani must have felt that everything she knew, everything she was familiar with – the water, the currents, the earth itself – was rising up against her.’

