As a real-world example, pick any extended conflict in the world: Israelis versus Palestinians, Hutus versus Tutsis, Bosnians versus Serbs, Sunni versus Shia. Climbing out on a limb here, I’d like to suggest that no living member of these groups is at fault for the anger that they feel toward each other, since the conflicts in question began many generations ago. But each individual today does bear some responsibility for continuing the conflict, because it’s possible for each person to change their concepts and therefore their behavior. No particular conflict is predetermined by evolution.
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