As May looked at more and more biological systems through the prism of simple chaotic models, he continued to see results that violated the standard intuition of practitioners. In epidemiology, for example, it was well known that epidemics tend to come in cycles, regular or irregular. Measles, polio, rubella—all rise and fall in frequency. May realized that the oscillations could be reproduced by a nonlinear model and he wondered what would happen if such a system received a sudden kick—a perturbation of the kind that might correspond to a program of inoculation. Naïve intuition suggests that
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On nonlinear reactions to vaccines
