Barry Grimes

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Many animals can experience unpleasant affect when another animal nearby is suffering. The first animal’s body budget is taxed by the second animal’s discomfort, so the first animal tries to fix the situation.* Even a rat will help another rat who is in distress, for example. Human infants can comfort another infant who is in distress. You don’t need emotion concepts for this ability, just a nervous system with interoception that produces affect.
How Emotions Are Made: The Secret Life of the Brain
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