Anatomical structures have no special status over behavioural ones, where ‘direct’ effects of genes are concerned. Genes are ‘really’ or ‘directly’ responsible only for proteins or other immediate biochemical effects. All other effects, whether on anatomical or behavioural phenotypes, are indirect. But the distinction between direct and indirect is vacuous. What matters in the Darwinian sense is that differences between genes are rendered as differences in phenotypes.
