‘You can observe lions in the field, but they’ll just lie there, and you can work with wolves for years and just see scats or hear howls,’ says hyena aficionado Kevin Theis. ‘But hyenas . . . there are greetings, reintroductions, dominance and submissive signalling. You’ll have young cubs trying to learn their place within the clan, immigrant males doing a run-through to see who’s there. Their social lives are incredibly more complex.’
