Doing genetic experiments with bees is quite a complicated business for various reasons. Worker bees themselves do not ordinarily reproduce, and so you have to cross a queen of one strain with a drone (= male) of the other, and then look at the behaviour of the daughter workers. This is what W. C. Rothenbuhler did. He found that all first-generation hybrid daughter hives were non-hygienic: the behaviour of their hygienic parent seemed to have been lost, although as things turned out the hygienic genes were still there but were recessive, like human genes for blue eyes. When Rothenbuhler
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