Empedocles calls them Love and Strife. These two principles, which we might think of as cosmic forces, are ultimately responsible for the formation of plants, animals, stars—the whole cosmos—out of the four roots (§§349, 355, 360). They play something like the role that Mind plays in the cosmology of Anaxagoras, except that, of course, in Anaxagoras Mind has no other force opposing it. The fact that Empedocles has two principles gives him the opportunity to put forward a grandiose and influential idea: his theory of cosmic cycles.
