In truth, Darwin knew, nature was one grand cacophonous battle—brutal, unyielding, and cruel. For if populations in the wild have such high rates of fertility that their size would increase exponentially if not constrained; if it is known that, excepting seasonal fluctuations, the size of populations remains stable over time; if Malthus was right, as he surely was, that the resources available to a species are limited—then it follows that there must be intense competition, or a struggle for existence, among the members of a species. And if no two members of a population are identical, and some
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