3 Or as Frazzetta himself observed, “I thus find it difficult to envision a smooth transition from a single maxilla to the divided condition seen in bolyerines.”4 Yet because the intermediate forms would not be viable, building a bolyerine jaw would require all the necessary parts—the jointed maxilla, the adjoining ligaments, and the necessary muscles and tissues—arising together. Yet the problem for neo-Darwinian theory, Frazzetta realized, extended well beyond the anatomical peculiarities of rare snakes. As a young evolutionary biology professor, he had studied complex features in a wide
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