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424 pages, Paperback
First published May 27, 2014
Crustose lichens, as the name suggests, form crustlike patches on the substrate. Some species have tiny lobes around the edges that can be somewhat ascending, and some are mostly adnate (flat), but they lack a lower cortex, so the tissues of the medulla are tightly bound to the substrate. The thallus can be thick or thin, continuous, cracked (rimose), divided into irregular sections (areolate), or consist of granules or rough warts (verrucose, or verruculose when the warts are minute).