The woeful inefficiency of knuckle walking is rarely a problem for chimpanzees deep in the forest, but it must have been a serious challenge for our missing-link ancestors about seven to nine million years ago. During this period of rapid climate change, the rain forest that covered much of Africa shrank and split into thousands of fragments interspersed with drier, open woodlands. For apes living in the depth of the rain forests, life went on as usual, but those at the margins of the forest must have faced a crisis. As woodlands replaced the forest, the fruits that dominated their diet became
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